REVISION OF THE ASIAN FOREST SCORPIONS (HETEROMETRINAE SIMON, 1879)

Heterometrus sp

Heterometrus glaucus (Thorell, 1876)
Heterometrus laevigatus (Thorell, 1876)
Heterometrus laoticus (Couzijn, 1981)
Heterometrus longimanus (Herbst, 1800)
Heterometrus petersii (Thorell, 1876)
Heterometrus silenus (Simon, 1884)
Heterometrus spinifer (Ehrenberg, 1828)
Heterometrus thorellii (Pocock, 1897)

Chersonesometrus sp

Chersonesometrus bastawadei (Prendini & Loria, 2020)
Chersonesometrus beccaloniae (Kovařík, 2004)
Chersonesometrus fulvipes (C. L. Koch, 1837)
Chersonesometrus hendersoni (Prendini & Loria, 2020)
Chersonesometrus madraspatensis (Pocock, 1900)
Chersonesometrus nathanorum (Prendini & Loria, 2020)
Chersonesometrus pelekomanus (Couzijn, 1981)
Chersonesometrus shivashankari (Prendini & Loria, 2020)
Chersonesometrus tristis (Henderson, 1919)
Chersonesometrus wroughtoni (Pocock, 1899)

Deccanometrus sp

Deccanometrus bengalensis (C. L. Koch, 1841)
Deccanometrus latimanus (Pocock, 1894)
Deccanometrus liurus (Pocock, 1897)
Deccanometrus obscurus (Couzijn, 1981)
Deccanometrus phipsoni (Pocock, 1893)
Deccanometrus ubicki (Kovarik, 2004)
Deccanometrus xanthopus (Pocock, 1897)

Gigantometrus sp

Gigantometrus swammerdami (Simon, 1872)
Gigantometrus titanicus (Couzijn, 1981)

 

Javanimetrus sp

Javanimetrus cyaneus (C. L. Koch, 1836)

Sahyadrimetrus sp

Sahyadrimetrus barberi (Pocock, 1900)
Sahyadrimetrus kanarensis (Pocock, 1900)
Sahyadrimetrus mathewi (Prendini & Loria, 2020)
Sahyadrimetrus rugosus (Couzijn, 1981)
Sahyadrimetrus scaber (Thorell, 1876)
Sahyadrimetrus tikaderi (Prendini & Loria, 2020)

Rugodentus keralaensis (Bastawade, Sureshan & Radhakrishnan, 2005)

Srilankametrus caesar (C. L. Koch, 1841)
Srilankametrus couzijni (Prendini & Loria, 2020)
Srilankametrus gravimanus (Pocock, 1894)
Srilankametrus indus (DeGeer, 1778)
Srilankametrus pococki (Prendini & Loria, 2020)
Srilankametrus serratus (Pocock, 1900)
Srilankametrus yaleensis (Kovarik, Ranawana, Jayarathne, Hoferek & Stahlavsky, 2019)

Srilankametrus serratus (Pocock, 1900), comb. nov; Srilankametrus yaleensis (Kovarik et al., 2019), comb. nov. Twenty-seven new synonyms are presented: Scorpio leioderma Dufour,
1856 = Sahyadrimetrus scaber (Thorell, 1876), syn. nov.; Palamnaeus costimanus var. B borneensis
Thorell, 1876 = Heterometrus longimanus (Herbst, 1800), syn. nov.; Palamnaeus liophysa ‘Thorell,
1888 = Heterometrus longimanus (Herbst, 1800), syn. nov.; Palamnaeus oatesii Pocock, 1900 = Het-
erometrus petersii (Thorell, 1876), syn. nov.; Palamnaeus swammerdami flavimanus Pocock, 1900 =
Gigantometrus swammerdami (Simon, 1872), syn. nov.; Heterometrus liophysa var. madoerensis Kop-
stein, 1921 = Heterometrus glaucus (Thorell, 1876), syn. nov.; Heterometrus laevifrons Roewer, 1943
= Heterometrus glaucus (Thorell, 1876), syn. nov.; Heterometrus (Chersonesometrus) granulomanus
Couzijn, 1981 = Srilankametrus caesar (C.L. Koch, 1841), syn. nov.; Heterometrus (Heterometrus)
liophysa separatus Couzijn, 1981 = Heterometrus glaucus (Thorell, 1876), syn. nov.; Heterometrus
(Heterometrus) liophysa spartanicus Couzijn, 1981 = Heterometrus glaucus (Thorell, 1876), syn. nov.;
Heterometrus (Heterometrus) longimanus bengkalitensis Couzijn, 1981 = Heterometrus longimanus
(Herbst, 1800), syn. nov.; Heterometrus (Heterometrus) longimanus marmoratus Couzijn, 1981 = Het-
erometrus longimanus (Herbst, 1800), syn. nov.; Heterometrus (Heterometrus) petersii mindanaensis
Couzijn, 1981 = Heterometrus silenus (Simon, 1884), syn. nov.; Heterometrus (Heterometrus) spinifer
solitarius Couzijn, 1981 = Heterometrus spinifer (Ehrenberg, 1828), syn. nov.; Heterometrus (Srilanka-
metrus) indus laevitensus Couzijn, 1981 = Srilankametrus indus (DeGeer, 1778), syn. nov.; Heterome-
trus (Heterometrus) keralaensis Tikader and Bastawade, 1983 = Sahyadrimetrus rugosus (Couzijn,
1981), syn. nov.; Heterometrus cimrmani Kovatik, 2004 = Heterometrus laevigatus (Thorell, 1876),
syn. nov.; Heterometrus mysorensis Kovarik, 2004 = Chersonesometrus tristis (Henderson, 1919), syn.
nov.; Heterometrus nepalensis Kovarik, 2004 = Deccanometrus bengalensis (Pocock, 1900), syn. nov.;
Heterometrus rolciki Kovatik, 2004 = Sahyadrimetrus scaber (Thorell, 1876), syn. nov.; Heterometrus
sejnai Kovatik, 2004 = Javanimetrus cyaneus (C.L. Koch, 1836), syn. nov.; Heterometrus tibetanus
Lourenco et al., 2005 = Deccanometrus bengalensis (Pocock, 1900), syn. nov.; Heterometrus liangi Zhu
and Yang, 2007 = Heterometrus silenus (Simon, 1884), syn. nov.; Heterometrus telanganaensis Javed
et al., 2010 = Deccanometrus xanthopus (Pocock, 1897), syn. nov.; Heterometrus atrascorpius Mirza
et al., 2012 = Chersonesometrus beccaloniae (Kovatik, 2004), syn. nov.; Heterometrus minotaurus
Pliskova et al., 2016 = Heterometrus laevigatus (Thorell, 1876), syn. nov.; Heterometrus bastawadei
Rossi, 2016 = Rugodentus keralaensis Bastawade et al., 2005, syn. nov. Another 25 synonyms by
previous authors are confirmed, for a total of 51 synonyms in subfamily Heterometrinae. Revised
diagnoses with comparative images, and a key and distributional atlas of the genera and species are
provided, along with a summary of available data for their ecology and conservation status, where
applicable.