Heterometrus

Heterometrus cimrmani (Kovařík, 2004)
Heterometrus glaucus (Thorell, 1876)
Heterometrus laevigatus (Thorell, 1876)
Heterometrus laoticus (Couzijn, 1981)
Heterometrus longimanus (Herbst, 1800)
Heterometrus petersii (Thorell, 1876)
Heterometrus silenus (Simon, 1884)
Heterometrus spinifer (Ehrenberg, 1828)
Heterometrus thorellii (Pocock, 1897)

Heterometrus cimrmani (Kovařík, 2004)

H. cimrmani, male from Trang (50) and female from Nakhon Si Thammarat (51), THNHM.

Figures 58–59. H. cimrmani from Trang.

Figures 62–65: Heterometrus cimrmani, in dorsal (62, 64) and ventral (63, 65) aspects; scale bar = 10 mm. Figures 62–63. Male from Trang,
THNHM. Figures 64–65. Female from Nakhon Si Thammarat, THNHM.

Figure 66. H. cimrmani from Trang.

Arm-span competition between males of Heterometrus cimrmani from Phattalung.\

Reference

Kawai, K., Unnahachote, T., Suttisatid, Y., & Tang, V. 2023 . А review of Heterometrus in Thailand (Scorpiones: Scorpionidae). Euscorpius, No. 373: 1-25.

Heterometrus glaucus (Thorell, 1876)

H.glaucus

Reference

SYSTEMATIC REVISION OF THE ASIAN FOREST SCORPIONS (HETEROMETRINAE SIMON, 1879),
REVISED SUPRAGENERIC CLASSIFICATION OF SCORPIONIDAE LATREILLE, 1802, AND REVALIDATION OF RUGODENTIDAE BASTAWADE ET AL., 2005
LORENZO PRENDINI
Scorpion Systematics Research Group,
Division of Invertebrate Zoology,
American Museum of Natural History
STEPHANIE F. LORIA
Scorpion Systematics Research Group,
Division of Invertebrate Zoology;
Richard Gilder Graduate School,
American Museum of Natural History

Heterometrus laevigatus (Thorell, 1876)

FIGURE 23. Heterometrus Ehrenberg, 1828, carapace, dorsal aspect. A, B. Heterometrus laevigatus (‘Thorell,
1876), comb. nov. et stat. rev., A. d (AMNH), Thale Ban, Thailand, B. 2 (AMNH), Khao Pu-Khao Ya, Thai-
land.

FIGURE 50, Heterometrus Ehrenberg, 1828, legs I-IV, tibiae, basitarsi and telotarsi, ventral aspect. A-D. Het-
erometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., 6 (AMNH), Thale Ban, Thailand.

D. Heterometrus laevigatus
(Thorell, 1876), comb. nov. et stat. rev., d (AMNH), Thale Ban, Thailand.

FIGURE 166. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev. d (AMNH), Thale Ban, Thai-
land, pedipalp patella: A. dorsal, B. retrolateral, and C. ventral aspects. Scale bar = 5 mm.

$image.png$ Figures 1–3: Heterometrus laevigatus, holotype subadult female (NMG 90); scale bar = 10mm. Figure 1. Dorsal aspect. Figure 2. Ventral
aspect. Figure 3. Label.

Figures 22–23. H. minotaurus, male (22)

H. minotaurus, male (32,42,52)

H. cimrmani from Trang. Figures 60–61.
H. minotaurus from Surat Thani.

Reference

А Review of Heterometrus in Thailand
(Scorpiones: Scorpionidae)
Kazusa Kawai, Thornthan Unnahachote,
Yossaphon Suttisatid and Victoria Tang
June 2023 — No. 373

Heterometrus laoticus (Couzijn, 1981)

FIGURE 171. Heterometrus laoticus Couzijn, 1981,
d (AMNH), Phahom Camp, Laos, pedipalp patella:
A. dorsal, B. retrolateral, and C. ventral aspects.
Scale bar = 5 mm.

A, B. Heterometrus laoticus Couzijn,
1981, A. d (AMNH), Phahom Camp, Laos, B. 2 (AMNH), Tham Pha In, Laos.

A, B. Heterometrus laoticus
Couzijn, 1981, A. ¢ (AMNH), Phahom Camp, Laos, B. 2 (AMNH), Tham Pha In, Laos.

A-D. Het-
erometrus laoticus Couzijn, 1981, d6 (AMNH), Phahom Camp, Laos.

A-D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev.,
metasoma and telson, ventral aspect.

Reference

Heterometrus longimanus (Herbst, 1800)

FIGURE 176. Heterometrus longimanus (Herbst, 1800), 6 (AMNH), Kuching, Malaysia, pedipalp patella: A.
dorsal, B. retrolateral, and C. ventral aspects. Scale bar = 5 mm.

FIGURE 177. Heterometrus longimanus (Herbst, 1800), pedipalp chela: A, B. retrodorsal, C. ventral, and D.
prolateral aspects. A. 2 (AMNH), B-D. ¢ (AMNH), Kuching, Malaysia. Scale bar = 5 mm.

Reference

Heterometrus petersii (Thorell, 1876)

FIGURE 158. Map of Southeast Asia, plotting known locality records of five species of Heterometrus Ehrenberg,
1828, based on material examined, and verified records from virtual museums and the literature: Heterometrus
laevigatus (Thorell, 1876), comb. nov. et stat. rev.; Heterometrus longimanus (Herbst, 1800); Heterometrus petersii
(Thorell, 1876); Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev.; Heterometrus thorellii (Pocock, 1892).

FIGURE 178. Heterometrus petersii (Thorell, 1876), holotype 2 (NMG 91), habitus: A. dorsal and B. ventral
aspects. Scale bar = 10 mm.

Reference

Heterometrus silenus (Simon, 1884)

Heterometrus silenus (part): Prendini & Loria, 2020: 279–
290, figs. 10, 24C, D, 51E–H, 70C, 72C, 158, 179–183,
table 2 (reference list until 2020).

C, D. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev. C. d, D. 2 (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale
bars = 2 mm.

C, D. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev. C. 6, D. 2 (AMNH), Ta Kou Mountain Nature Reserve, Viet-
nam. Scale bars = 2 mm.

E-H. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., d (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 2 mm.

E-H. Heterometrus longimanus (Herbst, 1800), ¢ (AMNH), Kuching, Malaysia. Scale bars = 2 mm.

C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., 6 (AMNH), Ta Kou Mountain Nature Reserve, Vietnam.

FIGURE 179. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., habitus, dorsal aspect: A. d
(AMNH), B. 2 (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 10 mm.

FIGURE 180. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., habitus, ventral aspect: A. d
(AMNH), B. 2 (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 10 mm.

FIGURE 181. Heterometrus silenus (Simon, 1884),comb. nov. et stat. rev., ¢6 (AMNH), Ta Kou Mountain Nature Reserve, Vietnam,pedipalp patella: Adorsal, B. retrolateral, and C. ventral aspects. Scale
bar = 5 mm.

FIGURE 182. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., pedipalp chela, retrodorsal aspect.
A. 2 (AMNH), B. d (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bar = 5 mm.

FIGURE 183. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., 6 (AMNH), Ta Kou Mountain
Nature Reserve, Vietnam, pedipalp chela: A. ventral and B. nprolateral aspects. Scale bar = 5 mm.

Figures 8–9. H. silenus, male from Chanthaburi (8) and female from Chonburi (9), TUPC.

Figures 16–17. H. silenus, male from Chanthaburi (16) and female from Chonburi (17), TUPC.

Figures 24–25. H. silenus, male from Chanthaburi (24) and female from Chonburi (25), TUPC.

Figures 34–35. H. silenus, male from Chanthaburi (34) and female from Chonburi (35), TUPC.

Figures 44–45. H. silenus, male from Chanthaburi (44) and female from Chonburi (45),
TUPC.

gures 54–55. H. silenus, male from Chanthaburi (54) and female from Chonburi (55), TUPC.

Figures 76–79: Heterometrus silenus, in dorsal (76, 78) and ventral (77, 79) aspects; scale bar = 10 mm. Figures 76–77. Male from Chanthaburi,
TUPC. Figures 78–79. Female from Chonburi, TUPC.

Reference

А Review of Heterometrus in Thailand
(Scorpiones: Scorpionidae)
Kazusa Kawai, Thornthan Unnahachote,
Yossaphon Suttisatid and Victoria Tang
June 2023 — No. 373

Heterometrus spinifer (Ehrenberg, 1828)

C, D. Heterometrus spinifer (Ehrenberg, 1828), C. 6 (AMNH), D. 2 (AMNH), Cameron Highlands,
Malaysia. Scale bars = 2 mm.

C, D. Heterometrus spinifer (Ehrenberg, 1828), C. d, D. 2 (AMNH),
Cameron Highlands, Malaysia. Scale bars = 2 mm.

E-H. Heterome-
trus spinifer (Ehrenberg, 1828), 6 (AMNH), Cameron Highlands, Malaysia. Scale bars = 2 mm.

FIGURE 186. Heterometrus spinifer (Ehrenberg,
1828), ¢ (AMNH), Cameron Highlands, Malaysia,
pedipalp patella: A. dorsal, B. retrolateral, and C.
ventral aspects. Scale bar = 5 mm,

Figures 28–29. H. spinifer, males from Yala (28) and Narathiwat (29), TUPC.

Figures 80–81: Heterometrus spinifer, in vivo habitus and dorsal aspect. Figure 80. Male from Yala, TUPC. Figure 81. Male from Narathiwat,
TUPC.

$image.png$ Figures 83: Arm-span competition between males of Heterometrus spinifer from Yala (right) and Narathiwat (left).

Reference

А Review of Heterometrus in Thailand
(Scorpiones: Scorpionidae)
Kazusa Kawai, Thornthan Unnahachote,
Yossaphon Suttisatid and Victoria Tang
June 2023 — No. 373

Heterometrus thorellii (Pocock, 1897)

E, F. Heterometrus thorellii (Pocock, 1892), E. 3
(AMNH), Mandalay, Myanmar, F. 2 (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. Scale bars = 2 mm.

E, FE. Heterometrus thorellii (Pocock, 1892), E. d (AMNH), Mandalay, Myanmar, F. 2 (AMNH), Pho Kyar
Elephant Camp, Yedashe, Myanmar. Scale bars = 2 mm.

A-D. Heterometrus thorellii (Pocock, 1892),
(AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar.

FIGURE 190. Heterometrus thorellii (Pocock, 1892), 2 (AMNH), Pho Kyar Elephant Camp, Yedashe, Myan-
mar, pedipalp patella: A. dorsal, B. retrolateral, and C. ventral aspects. Scale bar = 5 mm.

Reference